The Voyage Of The Beagle By Charles Darwin





































































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The belief that where large quadrupeds exist, the vegetation
must necessarily be luxuriant, is the more remarkable,
because the converse - Page 36
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The Belief That Where Large Quadrupeds Exist, The Vegetation Must Necessarily Be Luxuriant, Is The More Remarkable, Because The Converse Is Far From True.

Mr. Burchell observed to me that when entering Brazil, nothing struck him more forcibly than the splendour of the South American vegetation contrasted with that of South Africa, together with the absence of all large quadrupeds.

In his Travels, [6] he has suggested that the comparison of the respective weights (if there were sufficient data) of an equal number of the largest herbivorous quadrupeds of each country would be extremely curious. If we take on the one side, the elephant, [7] hippopotamus, giraffe, bos caffer, elan, certainly three, and probably five species of rhinoceros; and on the American side, two tapirs, the guanaco, three deer, the vicuna, peccari, capybara (after which we must choose from the monkeys to complete the number), and then place these two groups alongside each other, it is not easy to conceive ranks more disproportionate in size. After the above facts, we are compelled to conclude, against anterior probability, [8] that among the mammalia there exists no close relation between the bulk of the species, and the _quantity_ of the vegetation, in the countries which they inhabit.

With regard to the number of large quadrupeds, there certainly exists no quarter of the globe which will bear comparison with Southern Africa. After the different statements which have been given, the extremely desert character of that region will not be disputed. In the European division of the world, we must look back to the tertiary epochs, to find a condition of things among the mammalia, resembling that now existing at the Cape of Good Hope. Those tertiary epochs, which we are apt to consider as abounding to an astonishing degree with large animals, because we find the remains of many ages accumulated at certain spots, could hardly boast of more large quadrupeds than Southern Africa does at present. If we speculate on the condition of the vegetation during these epochs we are at least bound so far to consider existing analogies, as not to urge as absolutely necessary a luxuriant vegetation, when we see a state of things so totally different at the Cape of Good Hope.

We know [9] that the extreme regions of North America, many degrees beyond the limit where the ground at the depth of a few feet remains perpetually congealed, are covered by forests of large and tall trees. In a like manner, in Siberia, we have woods of birch, fir, aspen, and larch, growing in a latitude [10] (64 degs.) where the mean temperature of the air falls below the freezing point, and where the earth is so completely frozen, that the carcass of an animal embedded in it is perfectly preserved. With these facts we must grant, as far as _quantity alone_ of vegetation is concerned, that the great quadrupeds of the later tertiary epochs might, in most parts of Northern Europe and Asia, have lived on the spots where their remains are now found. I do not here speak of the kind of vegetation necessary for their support; because, as there is evidence of physical changes, and as the animals have become extinct, so may we suppose that the species of plants have likewise been changed.

These remarks, I may be permitted to add, directly bear on the case of the Siberian animals preserved in ice. The firm conviction of the necessity of a vegetation possessing a character of tropical luxuriance, to support such large animals, and the impossibility of reconciling this with the proximity of perpetual congelation, was one chief cause of the several theories of sudden revolutions of climate, and of overwhelming catastrophes, which were invented to account for their entombment. I am far from supposing that the climate has not changed since the period when those animals lived, which now lie buried in the ice. At present I only wish to show, that as far as _quantity_ of food _alone_ is concerned, the ancient rhinoceroses might have roamed over the _steppes_ of central Siberia (the northern parts probably being under water) even in their present condition, as well as the living rhinoceroses and elephants over the _Karros_ of Southern Africa.

I will now give an account of the habits of some of the more interesting birds which are common on the wild plains of Northern Patagonia: and first for the largest, or South American ostrich. The ordinary habits of the ostrich are familiar to every one. They live on vegetable matter, such as roots and grass; but at Bahia Blanca I have repeatedly seen three or four come down at low water to the extensive mud-banks which are then dry, for the sake, as the Gauchos say, of feeding on small fish. Although the ostrich in its habits is so shy, wary, and solitary, and although so fleet in its pace, it is caught without much difficulty by the Indian or Gaucho armed with the bolas. When several horsemen appear in a semicircle, it becomes confounded, and does not know which way to escape. They generally prefer running against the wind; yet at the first start they expand their wings, and like a vessel make all sail. On one fine hot day I saw several ostriches enter a bed of tall rushes, where they squatted concealed, till quite closely approached. It is not generally known that ostriches readily take to the water. Mr. King informs me that at the Bay of San Blas, and at Port Valdes in Patagonia, he saw these birds swimming several times from island to island. They ran into the water both when driven down to a point, and likewise of their own accord when not frightened: the distance crossed was about two hundred yards. When swimming, very little of their bodies appear above water; their necks are extended a little forward, and their progress is slow. On two occasions I saw some ostriches swimming across the Santa Cruz river, where its course was about four hundred yards wide, and the stream rapid.

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